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W. Carter Johnson photographed this Celastrus scandens while conducting research in the Garrison Reach of the Missouri River in North Dakota.
Celastrus scandens
Monarch butterfly larve in the Garrison Reach of the Missouri River in North Dakota.
Monarch larve
W. Carter Johnson photographs these Missouri black-billed cuckoo eggs in a nest while conducting research in the Garrison Reach of the Missouri River in North Dakota.
Missouri Black-billed cuckoo nest
Hummocky ground on North Dakota farm.
Hummocky Ground, North Dakota Farm
Glacial till in central North Dakota road cut.
Glacial Till Exposed at Road Cut
Pasque flower (Anemone patens) in North Dakota prairie.
Anemone patens
Reach stand # 4 near the Garrison dam on the Missouri River.
Research stand on the Missouri River
Research stand #8 near the Garrison dam on the Missouri River.
Research stand on the Missouri River
Research stand #6 on the Missouri River.
Research stand on the Missouri River
Research stand #3 on the Missouri River.
Research stand on the Missouri River
Fall Celastrus vegetation at research stand #11 near the Garrison dam on the Missouri River.
Fall Celatrus at research stand on the Missouri River
Mid-successional cottonwood forest with ash in the understory along the Missouri River at Garrison Reach in North Dakota.
Cottonwood forest at Garrison Reach, North Dakota
Large American elm trees at the Garrison Reach of the Missouri River in North Dakota. The truck in the background is the field vehicle used by Carter Johnson and Warren Keammerer.
Large American Elms at Garrison Reach, North Dakota
Garrison Reach of the Missouri River in North Dakota in the fall.
Garrison Reach of the Missouri River in North Dakota
Floodplain understory vegetation at Garrison Reach of the Missouri River in North Dakota.
Floodplain understory at Garrison Reach of the Missouri River, North Dakota
Painted woods at Garrison Reach of the Missouri River in North Dakota.
Painted woods at Garrison Reach of the Missouri River in North Dakota.
Trees flooded from the backup at Oahe Reservoir at Garrison Reach of the Missouri River in North Dakota.
Oahe Reservoir backup at Garrison Reach of the Missouri River, North Dakota
Trees flooded from the backup at Oahe Reservoir at Garrison Reach of the Missouri River in North Dakota.
Oahe Reservoir backup at Garrison Reach of the Missouri River, North Dakota
Surface water flooding of the Missouri River at Garrison Reach, south of Bismarck, North Dakota when the Oahe Reservoir was filled.
Oahe Reservoir backup at Garrison Reach of the Missouri River, North Dakota
J. Dodd and B. Lauenroth, fellow graduate students at North Dakota State University with W. Carter Johnson.
J. Dodd and B. Lauenroth, graduate students at NDSU
Buffalo grass (Buchloe dactyloides) with Japanese brome (Bromus japonicus) in Western North Dakota.
Bromus jap and Buchloe dacty
Scarlet globe mallow (Sphaeralcea coccinea) in western North Dakota.
Sphaeralcea coccinea
Scarlet globe mallow (Sphaeralcea coccinea) in western North Dakota.
Sphaeralcea coccinea
Big bluestem (Andropogon gerardii) at edge of floodplain forest along the Missouri River in North Dakota.
Andropogon gerardi
Cottonwood tree is 19.5 feet tall. In 2008 it was discovered that this tree and most other extremely large and old cottonwoods in this stand had died and fallen to the ground. This tree may have been present when Lewis and Clark passed by in the early 1800's. W. Carter Johnson's wife, Janet, is next to tree.
Large Cottonwood tree
Rolling field in central North Dakota.
Rolling Field
Prairie smoke (Geum triflorum) in North Dakota prairie.
Geum triflorum
W. Carter Johnson and Warren's research stands numbered 1-34.
Research stand
Floodplain forest research near the Garrison dam on the Missouri River.
Floodplain forest research on the Missouri River
Research stand #10 near the Garrison dam on the Missouri River.
Research stand on the Missouri River
Research stand #3 near the Garrison dam on the Missouri River.
Research stand on the Missouri River
Old cottonwood stand at Garrison Reach in North Dakota.
Old cottonwood stand at Garrison Reach, North Dakota
Dead trees in a clearing at Garrison Reach in North Dakota.
Clearing at Garrison Reach, North Dakota
Missouri River at Garrison Reach, North Dakota
Missouri River at Garrison Reach, North Dakota
Missouri River at Garrison Reach in North Dakota.
Missouri River at Garrison Reach, North Dakota
Emergent cottonwood trees at stand number 12 sampled by Carter Johnson during his doctoral research on the Missouri River floodplain at Garrison Reach.
Emergent cottonwood at Garrison Reach of the Missouri River, North Dakota
Trees eventually killed by the backup of the Missouri River at Garrison Reach, south of Bismarck, North Dakota when the Oahe Reservoir was filled.
Oahe Reservoir backup at Garrison Reach of the Missouri River, North Dakota
Cottonwood tree is 19.5 feet tall at research stand #26 in Garrison Reach on the Missouri River in North Dakota.
Large Cottonwood tree
Road cut exposing glacial till in central North Dakota.
Road Cut Exposing Glacial Till
Ancient shoreline marked by exposed rocks in North Dakota.
Ancient Shoreline Marked by Exposed Rocks
Glacial till in central North Dakota road cut.
Glacial Till, Central North Dakota
Pasque flower (Anemone patens) in North Dakota prairie.
Anemone patens
Floodplain forest research near the Garrison dam on the Missouri River.
Floodplain forest research on the Missouri River
Early successional forest dominated by cottonwood, about 20 years old, at Garrison Reach near the Missouri River in North Dakota.
Young cottonwood stand at Garrison Reach, North Dakota
Old cottonwood stand at Garrison Reach in North Dakota.
Old cottonwood stand at Garrison Reach, North Dakota
River bank on the Missouri at Garrison Reach, North Dakota
River bank on the Missouri at Garrison Reach, North Dakota
Dense canopy of later successional stage cottonwood forests near stand #12 in the floodplain at Garrison Reach of the Missouri River in North Dakota.
Emergent cottonwood at Garrison Reach of the Missouri River, North Dakota
Water backup of the Missouri River at Garrison Reach, south of Bismarck, North Dakota when the Oahe Reservoir was filled.
Oahe Reservoir backup at Garrison Reach of the Missouri River, North Dakota
Warren R. Keammerer standing in melilotus at stand #8 in Garrison Reach in North Dakota.
Warren Keammerer standing in Melilotus
W. Carter Johnson photographed this Missouri grape fern during his research. It is found in the rich, late successional forests along the Missouri River.
Missouri grape fern
W. Carter Johnson photographed this Missouri grape fern during his research. It is found in the rich, late successional forests along the Missouri River.
Missouri grape fern
Viola pennsylvanica in central North Dakota.
Polygala alba
Cicada in the Garrison Reach of the Missouri River in North Dakota.
Cicada
Prickly Pear cactus (Opuntia polyacantha) in western North Dakota.
Opuntia polyacantha
Prickly Pear cactus (Opuntia polyacantha) in western North Dakota.
Opuntia polyacantha
Indian breadroot (Psoralea esculenta) in western North Dakota. This plant was an important starch food plant for Native Americana
Psoralea esculenta
Penstemon grandiflorus in the Garrison Reach of the Missouri River in North Dakota.
Penstemon grandiflorus
Sunset on the Missouri River at the Garrison Reach in North Dakota.
Sunset on Missouri River
Effect of grazing on North Dakota farm.
North Dakota Farm Effects of Grazing
Research stand #2 destroyed by backup flooding of Oahe Reservoir on the Missouri River.
Research stand on the Missouri River
Reach stand # 4 near the Garrison dam on the Missouri River.
Research stand on the Missouri River
Research stand on the Missouri River
Research stand on the Missouri River
Research stand #1 near the Garrison dam on the Missouri River.
Research stand on the Missouri River
Research stand #6 near the Garrison dam on the Missouri River.
Research stand on the Missouri River
Vegetation growing on old cottonwood tree at Garrison Reach in North Dakota.
Vegetation on old cottonwood at Garrison Reach, North Dakota
Emergent cottonwood trees at Garrison Reach of the Missouri River in North Dakota. This image clearly shows how much taller old cottonwood trees are compared to the later successional stage. This exposure to storms causes wind damage and sometimes tree fall to emergent cottonwoods.
Emergent cottonwood at Garrison Reach of the Missouri River, North Dakota
Flooded forests at Garrison Reach of the Missouri River in North Dakota when Oahe Reservoir was filled. Surface water and groundwater levels had been increasing south of Bismarck due to sedimentation and flow backup. These trees photographed in 1969 most certainly died soon after.
Oahe Reservoir backup at Garrison Reach of the Missouri River, North Dakota
Water backup of the Missouri River at Garrison Reach, south of Bismarck, North Dakota when the Oahe Reservoir was filled.
Oahe Reservoir backup at Garrison Reach of the Missouri River, North Dakota
Water backup of the Missouri River at Garrison Reach, south of Bismarck, North Dakota when the Oahe Reservoir was filled.
Oahe Reservoir backup at Garrison Reach of the Missouri River, North Dakota
Warren R. Keammerer, who was Carter's graduate student colleague, working on the Missouri River forests at Painted Woods Lake in the Garrison Reach in North Dakota.
Warren Keammerer at Painted Woods Lake
Warren R. Keammerer is changing the tire of field vehicle used in river research.
Warren Keammerer changing a flat tire
Old cottonwood being recycled by decomposers in western North Dakota
Vegetation on fallen cottonwood
Spanish bayonet (Yucca glauca) in western North Dakota.
Yucca glauca
Missouri River break north of Bismarck, North Dakota on the east side of river.
Missouri River breaks
Janet Johnson stand next to the largest green ash tree sampled in W. Carter Johnson's research at #26 in the Garrison Reach of the Missouri River in North Dakota.
Large ash tree
Viola pensylvanica in North Dakota.
Viola pensylvanica
Ambrosia trifida in the Missouri River Valley in North Dakota.
Ambrosia trifida
Needle-and-thread grass (Stipa comata) in western North Dakota.
Stipa comata
Dead myrtle warbler in the Garrison Reach of the Missouri River in North Dakota.
Dead myrtle warbler
View from the top of the Prairie Coteau on the Dakota border.
Prairie Coteau
Uvularia grandiflora vegetation in North Dakota.
Uvularia grandiflora
Marsh on the floodplain at the Garrison Reach of the Missouri River in North Dakota.
Floodplain marsh at Garrison Reach, North Dakota
Expansive floodplain forests at the Garrison Reach of the Missouri River.
Floodplain forests on the Missouri River
Garrison Reach of the Missouri River in North Dakota in the fall.
Missouri River at Garrison Reach, North Dakota
Garrison Reach of the Missouri River in North Dakota in the fall.
Missouri River at Garrison Reach, North Dakota
Missouri River breaks on the western side of the Garrison Reach on the Missouri River in North Dakota.
Missouri River at Garrison Reach, North Dakota
Hayland near the Garrison Reach of the Missouri River in North Dakota.
Hayland near the Missouri River
Gaura coccinea at Garrison Reach on the Missouri River in North Dakota.
Gaura coccinea
Ratibida columnifera at Garrison Reach on the Missouri River in North Dakota.
Ratibida columnifera
Beaver dam in woody draw Cross Ranch in North Dakota.
Beaver dam
Mixed prairie storm in western North Dakota.
Mixed prairie storm
Desmodium canadense in western North Dakota.
Desmodium canadense
Glyceria grandis in western North Dakota
Glyceria grandis
Cornus stolonifers in Garrison Reach on the Missouri River in North Dakota.
Cornus stolonifers
Beaver dam in woody draw Cross Ranch in North Dakota. Photograph was taken before The Nature Conservancy purchase of the former Gaines Ranch.
Beaver dam
Asclepias verticillata in the Missouri River Valley in North Dakota.
Asclepias verticillata
W. Carter Johnson sitting on rock on Blossom Butte north of Bismarck, North Dakota on east side of the Missouri River. O. A. Stevens' book (green cover) is laying on the rock beside him.
W. Carter Johnson on Blossom Butte
Penstemon albidus in the Garrison Reach of the Missouri River in North Dakota.
Penstemon albidus
Penstemon gracilis in the Garrison Reach of the Missouri River in North Dakota.
Penstemon gracilis
Prairie Coteau stream in North Dakota.
Prairie Coteau stream
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